Daniel Pauly and Johannes Müller.

Pauly and Müller’s Breathing Water in a Warming World available on open access

Daniel Pauly and Johannes Müller.

Daniel Pauly (left) and Johannes Müller (right).

In a world where the dramatic effects of global warming on land dominate news headlines, there is a tendency to overlook the massive impacts of warming and deoxygenation on oceans and freshwater ecosystems. However, the effects of climate change on the species that inhabit these ecosystems are, in some respects, more drastic than the challenges faced by terrestrial animals.

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Orange-dotted grouper swimming and breathing in the ocean

New report sheds light on how fish grow in a warming, low-oxygen world

Orange-dotted grouper swimming and breathing in the ocean

Grouper. Image created with Adobe Firefly.

Aquatic animals that breathe through gills — including most fish and many invertebrates — are the backbone of life in oceans, lakes and rivers. They support biodiversity, shape food webs and sustain fisheries that feed millions of people worldwide. Understanding how these animals grow, reproduce and survive is therefore essential to understanding how aquatic ecosystems work — and how they continue to support human societies.

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Helostoma temminckii or kissing gourami with its mouth open.

Animal-welfare models fail to account for fish’s need for oxygen

Helostoma temminckii or kissing gourami with its mouth open.

Helostoma temminckii or kissing gourami. Image by Jörn, Wikimedia Commons.


A new essay published in Issues in Science and Technology argues that current animal welfare science and policy frameworks overlook a fundamental aspect of the lives of fish and other aquatic “water-breathing” species — and calls for a shift in how governments, researchers, and industry assess humane treatment in aquaculture, research, commercial fisheries, and in the wild.

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New GOLT book to be released in 2026

Breathing water in a warming world cover

A new book focused on the principles and applications of the Gill-Oxygen Limitation Theory (GOLT) is scheduled for publication in March 2026.

Co-authored by the Sea Around Us PI, Dr. Daniel Pauly, and Dr. Johannes Müller, assistant professor at the Leiden University Centre for the Arts in Society, Breathing Water in a Warming World presents a theoretical framework for explaining how warming waters and deoxygenation affect the growth and reproduction of fish and other water-breathing animals.

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